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The upgrades are a handy thing in this game, as they are pretty useful and easy to get. From the Memory Select screen, choose "Upgrades" and you'll be brought to a new menu. You can also watch the total sum of Bronze, Silver and Gold Templar Coins that you've collected. Each upgrade costs a certain amount of points. You get these points from collecting the aforementioned Templar Coins. Here's a list of the upgrades that are available:

Upgrade Cost Description
Hide Speed 50 points Altair becomes anonymous more quickly after hiding
Synchronization Rate 200 points Synchronization Rate Increase
Synchronization Strength 25 points 5 Synchronization Bars
50 Points 6 Synchronization Bars
75 Points 7 Synchronization Bars
100 Points 8 Synchronization Bars
125 Points 9 Synchronization Bars
Throwing Knife Maximum 100 Points 4 Throwing Knives
100 Points 5 Throwing Knives
Automatic Block 50 points Block Automatically
Critical Hits 75 Points 5% chance to critical hit
75 Points 10% chance to critical hit
75 Points 15% chance to critical hit
Upgrade Weapon Attack Damage 200 Points Level 1 increase
250 Points Level 2 increase
300 Points Level 3 increase

Note that after upgrading necessary skills you'll need to hit the "Confirm" option in order to store the changes you have made, and then exit to the Memory Select screen.