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Many of the game's actions are context sensitive. That means that the Action button performs differently depending on where you are and what you are trying to do. The same goes for the Command Ashley button.
PlayStation 2
GameCube
Wii
Action
Wii Remote & Nunchuk
Classic Controller
,
,
,
Move or turn, navigate menus
Move the camera
Pause the game and bring up the Options menu. skip cutscene
Action button (context sensitive)
Double-tap to kick open doors
,
Cancel/Hold down to run
,
Open inventory screen
Give commands to Ashley (context sensitive)
,
Open map
Pull out knife (hold and press the Action button to use it)
In the Wii version, you can just shake the Wii Remote for a quick knife attack
Equip weapon (hold and use Move control to aim, then press the Action button to fire/throw)