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|Toa Jaller||Toa Jaller can walk over lava, and after upgrading he can burn down passage-blocking vines. His Fireblast weapons are rapid-fire and are good at short- to mid-range, but spread out and become inaccurate at long range. Jaller's weapons include Toa Tahu's Fire Sword, Toa Vakama's Launcher, and his own Energized Flame Sword.|
|Toa Nuparu||Toa Nuparu is the slowest Toa but is quite powerful. He can climb certain walls and cliff faces, and after upgrading can dig up treasures. His Earthburst weapons fire grenade-like energy balls with a large blast radius that are slow to reload, but can be launched around corners and detonated manually. Nuparu's weapons include Toa Onua's Claw, Toa Whenua's Earthshock Drill, and his own Laser Drill.|
|Toa Hahli||Toa Hahli can cross streams and rivers, and after upgrading can manipulate blue water-based constractions. She has medium speed, but is stronger and faster when in water. Her long-range Watershock weapons don't have as much power, but as long as they are firing they cause steady damage. Hahli's weapons include Toa Gali's Hook, Toa Nokama's Hydro Blade, and her own Laser Harpoon.|
|Toa Hewkii||Toa Hewkii can assemble normal constractions from the various LEGO pieces found scattered in each level, and can upgrade this to the ability to collapse certain walls. Like Nuparu, he is slow but very strong. His Stonesmash weapons act like rocket launchers, and are slow to reload but have a large blast radius. Hewkii's weapons include the "Po-Koro" (based on Toa Pohatu's hands), Toa Onewa's Proto Pitons, and his own Laser Axe.|
|Toa Kongu||Toa Kongu is the second-fastest toa (Jaller's the fastest). Kongu can leap certain gaps marked by green landing pads, and can gain the ability to use green wind-based constractions. He has medium strength. His Airshot weapons are of medium strength, but grow weaker over long distances, as they possess a function similar to that of a shotgun. Kongu's weapons include Toa Lewa's Air Axe, Toa Matau's Aero Slicer, and his own Laser Crossbow.|
|Toa Matoro||Toa Matoro has the ability to zoom in towards targets, and some faraway white targets require the close-up view; he can also upgrade his ability to freeze water in places to make pathways. He is one of the slowest among the Toa, but has one of the strongest weapons available, able to kill some enemies in one hit. Toa Matoro's Icecrack weapons are similar to sniper rifles, with the ability to shoot lethal blasts at the expense of recharge time. Matoro's weapons include Toa Kopaka's Ice Sword, Toa Nuju's Crystal Spike, and his own Energized Ice Sword.|
In console versions of the game, each Piraka can also be played after they are defeated, though only in their own regions (for example, Vezok can only be played in the "Vezok's Coastline" levels); and they can activate special Piraka constractions. Once the game is completed, the individual Piraka are replaced by Vezon, who can activate not only Piraka constractions but black constractions found in each level.
In the Game Boy Advance version of the game, the six Toa Mata are playable instead of the Piraka, in both their original and transformed Toa Nuva incarnations. Later on, if you go back you can find the Toa Inika Masks and play as the Toa Inika.
Common enemies, which are encountered in each level, include each of the six breeds of Bohrok, Visorak, and Vahki. If the player collects all of the silver canisters in a level, they unlock one of the breeds of a common enemy, which can be viewed between levels.
In addition to the seven Piraka, there are 12 bosses which are unlocked for viewing after they are defeated: each of the six Rahkshi (Guurahk, Korahk, Lerahk, Panrahk, Turahk, and Vorahk), Roodaka, Sidorak, Nidhiki, Krekka, Brutaka, and Axonn.
Players also encounter Balta, a Matoran who lives on Voya Nui. In the game, he speaks in the introductory movie and runs the Shop where the player can purchase items and upgrades. He has Piruk's claws as hands in the intro, but in the upgrade shop, he is seen with his katanas.